73.320 Agrotis clavis (Heart & Club)
ws: 35-40mm (MBGBI9), fw: 14-18mm (Waring & Townsend), 13.1-17.0mm (my specimens); Jun-Aug; various herbaceous plants; common in S.England, more local from the Midlands north into Scotland.
ID: Ground colour ranges from pale straw to black; oval, kidney and dart marks prominent. Similar to A.exclamationis (Heart & Dart) and A.segetum (Turnip Moth) which have an equally variable range of ground colour. A.exclamationis can usually be distinguished by its prominent black collar (patagia) and in the male by the antennae lacking pectinations visible to the naked eye - males of A.segetum and A.clavis having antennae which are visibly pectinate in the basal half, and both sexes usually having a much less well-marked collar. The dart mark of A.segetum is usually dark outlined, while that of A.exclamationis and A.clavis is solidly dark. The forewing of A.clavis generally has a smaller length:breadth ratio than that of A.segetum or A.exclamationis, giving it a less elongate appearance; its dart mark is usually shorter and blunter; and its forewing is often clouded or flecked with brown - the forewing generally being plain in A.exclamationis.
Genital differences between A.segetum/clavis/exclamationis are covered by the Difficult Species Guide
See A.segetum/clavis/exclamationis for a direct comparison of these three species
Male genitalia: The 3 species are separated on features of the harpe. This is nearly straight in A.exclamationis; long and smoothly curved in A.clavis; stout and tightly curved, hook-like in A.segetum.
NB the harpe of §7 looks quite like A.segetum and I would have identified it as this species if it did not have a brownish-fuscous hindwing. I will leave it here until I have some reliable A.segetum material to compare. Moth dissection shows 2 dissections of A.segetum and of A.clavis. Neither of the A.clavis dissection have a harpe as broad or curved as §7; one of the dissections of A.segetum has a tighter curve than §7 but the other looks very similar to §7. Can A.segetum have a brownish-fuscous hindwing? do the harpes have a range of variation that overlaps?
Female genitalia: In A.segetum S8 has large rounded processes on either side of the ostium, which is not cupped, and the corpus bursae has a single linear signum. In A.clavis and A.exclamationis the rounded processes are absent, the ostium is cupped and there are no signa in the corpus bursae. These two differ in features of the appendix bursae which is shorter in A.clavis.
Genital differences between A.segetum/clavis/exclamationis are covered by the Difficult Species Guide
See A.segetum/clavis/exclamationis for a direct comparison of these three species
Male genitalia: The 3 species are separated on features of the harpe. This is nearly straight in A.exclamationis; long and smoothly curved in A.clavis; stout and tightly curved, hook-like in A.segetum.
NB the harpe of §7 looks quite like A.segetum and I would have identified it as this species if it did not have a brownish-fuscous hindwing. I will leave it here until I have some reliable A.segetum material to compare. Moth dissection shows 2 dissections of A.segetum and of A.clavis. Neither of the A.clavis dissection have a harpe as broad or curved as §7; one of the dissections of A.segetum has a tighter curve than §7 but the other looks very similar to §7. Can A.segetum have a brownish-fuscous hindwing? do the harpes have a range of variation that overlaps?
Female genitalia: In A.segetum S8 has large rounded processes on either side of the ostium, which is not cupped, and the corpus bursae has a single linear signum. In A.clavis and A.exclamationis the rounded processes are absent, the ostium is cupped and there are no signa in the corpus bursae. These two differ in features of the appendix bursae which is shorter in A.clavis.
§1 New Forest, Hampshire; 28/06/2009; female; fw 17.0mm
§2 Ham Fen, Kent; 15/07/2010; female; fw 16.2mm §3 Ham Fen, Kent; 15/07/2010; female; fw 16.4mm §4 Ham Fen, Kent; 15/07/2010; female; fw 15.4mm §5 Ham Fen, Kent; 15/07/2010; female; fw 16.5mm §6 Folkestone Warren, Kent; 03/07/2013; male; fw 15.5mm §7 Snettisham, Norfolk; 05/07/2013; male; fw 16.5mm §8 Snettisham, Norfolk; 12/06/2014; male; fw 16.0mm §9 Portland Bill, Dorset; 20/06/2015; male; fw 14.4mm All images © Chris Lewis |
§10 New Forest, Hampshire; 14/07/2019; male; fw 13.1mm
§11 Westcliff-on-sea, Essex; 24/06/2024; female; fw 17.7mm §12 Heydon, Cambridgeshire; 26/06/2024; male; fw 15.6mm |
Page published 19/01/2017 (§1-9) | §10 added 27/02/2020 | §11 added 21/08/2024 | §12 added 06/10/2024