Suborder:Glossata Infraorder:Heteroneura Division:Ditrysia Section:Cossina Subsection: Cossina Superfamily:Tortricoidea
Refs:
MBGBI5.1 Tortricinae & Chlidanotinae; MBGBI 5.2 Olethreutinae, Hancock, Bland & Razowski, 2014
British tortricoid moths, Bradley Tremewan & Smith, Cochylidae and Tortricidae:Tortricinae 1973 and Tortricidae:Olethreutinae 1979 - (BTM)
The genitalia of the group tortricidae of the lepidoptera of the British Isles, Pierce & Metcalfe, 1922
Field Guide to the micro moths of Great Britain and Ireland, Sterling & Parsons, 2012
Bird-dropping tortrix moths of the British Isles, Clifton & Wheeler, 2011
Wikipedia: List of Tortricidae Genera
Forewings broad with distinct tornal angle; Hindwings at least as broad as forewings
Male genitalia: uncus poorly developed, gnathos usually poorly developed, socii large and prominent; aedeagus curved and hinged with the juxta (pistol-like)
Key (MBGBI2): Cilia of hindwing shorter than breadth of hindwing > frenulum present > proboscis not scaled > ocelli present > hindtibia not long-haired > S2 of labial palp thicker than S3 > tarsi not spined beneath.
The RES Checklist recognises three subfamilies: Chlidanotinae, Tortricinae and Olethreutinae
Cochylini, now a tribe of Tortricinae, was considered as a separate family Chochylidae in BTM which also considers Chlidanotinae within Tortricinae.
Separation of subfamilies within BTM is based on venation:
Cochylinae: Forewing V1c obsolete, origin of V2 from distal 1/4 of discal cell
Tortricinae: Forewing V1c present, origin of V2 proximal to distal 1/4 of discal cell, hindwing without a cubital pecten on upper surface
Olethreutinae: Forewing V1c present, origin of V2 proximal to distal 1/4 of discal cell, hindwing with a cubital pecten on upper surface
(I have not yet found any information justifying the separation of Chlidanotinae and since Cochlylini is now placed within Tortricinae the above does not provide a clear justification for the current subfamily arrangement)
Oleuthreutinae can generally be recognised by the presence of costal strigulae - a pattern of light and dark marks, some of which may be extended as striae directed towards the subterminal region.
Pierce and Metcalfe divided the Tortricidae into 9 groups on characteristics of the female genitalia, almost entirely on the form of the signum/signa. These groups are mostly but not entirely coincident with the subfamilies and tribes considered here.
Most tortrices (tortricids, tortrixes) do not have widely accepted English names. Recently some English names have been proposed and are being used by Norfolk Moths and Suffolk Moths, I have adopted these as well.
Refs:
MBGBI5.1 Tortricinae & Chlidanotinae; MBGBI 5.2 Olethreutinae, Hancock, Bland & Razowski, 2014
British tortricoid moths, Bradley Tremewan & Smith, Cochylidae and Tortricidae:Tortricinae 1973 and Tortricidae:Olethreutinae 1979 - (BTM)
The genitalia of the group tortricidae of the lepidoptera of the British Isles, Pierce & Metcalfe, 1922
Field Guide to the micro moths of Great Britain and Ireland, Sterling & Parsons, 2012
Bird-dropping tortrix moths of the British Isles, Clifton & Wheeler, 2011
Wikipedia: List of Tortricidae Genera
Forewings broad with distinct tornal angle; Hindwings at least as broad as forewings
Male genitalia: uncus poorly developed, gnathos usually poorly developed, socii large and prominent; aedeagus curved and hinged with the juxta (pistol-like)
Key (MBGBI2): Cilia of hindwing shorter than breadth of hindwing > frenulum present > proboscis not scaled > ocelli present > hindtibia not long-haired > S2 of labial palp thicker than S3 > tarsi not spined beneath.
The RES Checklist recognises three subfamilies: Chlidanotinae, Tortricinae and Olethreutinae
Cochylini, now a tribe of Tortricinae, was considered as a separate family Chochylidae in BTM which also considers Chlidanotinae within Tortricinae.
Separation of subfamilies within BTM is based on venation:
Cochylinae: Forewing V1c obsolete, origin of V2 from distal 1/4 of discal cell
Tortricinae: Forewing V1c present, origin of V2 proximal to distal 1/4 of discal cell, hindwing without a cubital pecten on upper surface
Olethreutinae: Forewing V1c present, origin of V2 proximal to distal 1/4 of discal cell, hindwing with a cubital pecten on upper surface
(I have not yet found any information justifying the separation of Chlidanotinae and since Cochlylini is now placed within Tortricinae the above does not provide a clear justification for the current subfamily arrangement)
Oleuthreutinae can generally be recognised by the presence of costal strigulae - a pattern of light and dark marks, some of which may be extended as striae directed towards the subterminal region.
Pierce and Metcalfe divided the Tortricidae into 9 groups on characteristics of the female genitalia, almost entirely on the form of the signum/signa. These groups are mostly but not entirely coincident with the subfamilies and tribes considered here.
Most tortrices (tortricids, tortrixes) do not have widely accepted English names. Recently some English names have been proposed and are being used by Norfolk Moths and Suffolk Moths, I have adopted these as well.
Subfamily: Chlidanotinae (2G 2S)
Genera: Olindia, Isotrias
Genera: Olindia, Isotrias
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Subfamily: Tortricinae (7T 46G 140S[3EX])
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Tribe: Sparganothini (1G 1S)
Genus: Sparganothis 
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Tribe: Ramapesiini (7G 7S)
Genera: Ditula, Epagoge, Paramesia, Periclepsis, Philedone, Capua, Philedonides 
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Tribe: Archipini (15G 31S[2EX])
Genera: Archips (6S[1EX]), Choristoneura (3S[1EX]), Argyrotaenia, Ptycholomoides, Ptycholoma, Pandemis (5S), Syndemis, Lozotaenia, Cacoecimorpha, Aphelia (3S), Dichella, Clepsis (4S), Epiphyas, Lozotaenioides, Adoxophyes 
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Tribe: Cnephasiini (5G 15S[1EX])
Genera: Neosphaleroptera, Exapate, Tortricodes, Eana (4S[1EX]), Cnephasia (9S) 
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Tribe: Tortricini (4G 32S)
Genera: Spatalistis, Tortrix, Aleimma, Acleris (29S) 
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Tribe: Euliini (2G 2S)
Genera: Eulia, Pseudargyrotoza 
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Cochylini (12G 51S)
Genera: Phtheochroa (4S), Hysterophora, Cochylimorpha (2S), Phalonidia (5S), Gynnidomorpha (5S), Agapeta (2S), Eupoecilia (2S), Commophila,
Aethes (15S), Cochylidia (4S), Cochylis (8S), Falseuncaria (2S)
Genera: Phtheochroa (4S), Hysterophora, Cochylimorpha (2S), Phalonidia (5S), Gynnidomorpha (5S), Agapeta (2S), Eupoecilia (2S), Commophila,
Aethes (15S), Cochylidia (4S), Cochylis (8S), Falseuncaria (2S)
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Subfamily: Olethreutinae (7T 46G[2EX] 240S[5EX,1IR])
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Tribe: Olethreutini (13G[1EX] 38S[2EX])
Genera: Eudemis (2S), Pseudosciaphila, Apotomis (8S), Orthotaenia, Hedya (5S), Celypha (10S[1EX]), Phiaris (5S), Pristerognatha [1EX], Cymolomia, Argyroploce, Stictea, Olethreutes, Piniphila 
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Tribe: Lobesiini (1G 5S)
Genus: Lobesia (5S) 
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Endotheniini (1G 8S)
Genus: Endothenia (8S) 
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Tribe: Bactrini
Genus: Bactra (5S[1IR]) 
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Tribe: Enarmoniini (3G+1 19S+1)
Genera: Eucosmomorpha, Enarmonia, Tetramoera, Ancylis (17S) 
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Tribe: Eucosmini (20G[1EX] 91S[2EX])
Genera: Eriopsila, Thiodia, Rhopobota (4S), Spilonota (2S), Acroclita, Gibberifera (1EX), Epinotia (29S), Zeiraphera (4S), Crocidosema, Phaneta, Pelochrista, Eucosma (14S), Gypsonoma (6S[1EX]) Epiblema (8S), Notocelia (7S), Pseudococcyx (2S), Retinia, Gravitarmata, Clavigesta (2S), Rhyacionia (4S) 
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Grapholitini (7G 74S[1EX])
Genera: Dichrorampha (15S), Cydia (21S[1EX]), Lathronympha, Selania, Grapholita (13S), Pammene (21S), Strophodera (2S)
Genera: Dichrorampha (15S), Cydia (21S[1EX]), Lathronympha, Selania, Grapholita (13S), Pammene (21S), Strophodera (2S)
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