Subfamily: Olethreutinae
Tribe: Eucosmini (20G[1EX] 91S[2EX])
The genera included here are those of Pierce and Metcalfe's 'Epiblemidii' and 'Ephippiphoridii' characterised by the pair of thorn-like signa in the bursa copulatrix. The reason for considering the genera included in 'Ephippiphoridii' (Rhopobota, Spilonota, Gypsonoma, Gibberifera, Rhyacionia, Clavigesta) as distinct from 'Epibelmidii' is not clear. P&M consider the signa of 'Epiblemidii' to be 'pilleate' which they define in the glossary as "in the form of a Greek cap"; and those of 'Ephippiphoridii' to be 2 horns. Further confusion arises from the placement in 'Ephippiphoridii' of Eucosmomorpha now considered in Tribe: Enarmoniini and all the genera now placed in Tribe: Grapholitini except Dichrorampha.
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Eriopsela
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Thiodia
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Rhopobota (4S)
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218 Eriopsela quadrana
ws: 12-16mm; May-Jun; goldenrod (Solidago virgaurea); NS-B throughout GB |
219 Thiodia citrana
(Lemon Bell) 
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220 Rhopobota myrtillana
(Bilberry Bell) 
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221 Rhopobota stagnana
(Downland Bell) m11-18 f10-14mm; bivoltine Apr-Jun Aug-Sep; small/devil's-bit scabious (Scabiosa columbaria, Succisa pratensis); NS-B damp woodland and downs throughout GB |
222 Rhopobota ustomaculana
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223 Rhopobota naevana
(Holly Tortrix) 
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Spilonota (2S)
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Acroclita
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224 Spilonota ocellana
(Bud Moth) 
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225 Spilonota laricana
(Larch-bud Moth) 
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226 Acroclita subsequana
(Coast Spurge Bell) 
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Gibberifera (EX)
227 Gibberifera simplana
(Least Bell) Formerly in woodland in Essex, Kent & E.Sussex; now extinct in GB |
Epinotia (28S)
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228 Epinotia sordidana
(Brown Alder Bell) 
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229 Epinotia caprana
(Large Sallow Bell) 
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230 Epinotia trigonella
(White-blotch Bell) 
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231 Epinotia brunnichana
(Large Birch Bell) 
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232 Epinotia maculana
(Dark Aspen Bell) ws: 17-23mm; Aug-Oct; aspen (Populus tremula); NS-B woodland with aspen throughout GB |
233 Epinotia solandriana
(Variable Bell) 
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234 Epinotia abbreviana
(Brown Elm Bell) 
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235 Epinotia nemorivaga
ws: 10-12mm; Jun-Jul; bearberry (Arctostaphylos uva-ursi); NS-A Scottish highlands |
236 Epinotia granitana
(Scarce Spruce Bell) Rare migrant or accidnetal import, single British record 2007, Elevedon Forest |
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237 Epinotia signatana
(Black-brindled Bell) 
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238 Epinotia cruciana
(Willow Tortrix) 
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239 Epinotia mercuriana
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240 Epinotia immundana
(Common Birch Bell) 
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241 Epinotia crenana
ws:13-15mm; Jul-Aug; willows (Salix spp); RDB N.scotland eg Rothiemurchus, Strathspey, Skye, Rannoch |
242 Epinotia nanana
(Small Spruce Bell) 
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243 Epinotia demarniana
(Birch Bell) 
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244 Epinotia subocellana
(White Sallow Bell) 
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245 Epinotia tetraquetrana
(Square-barred Bell) 
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246 Epinotia pygmaeana
(Dingy Spruce Bell) 
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247 Epinotia subsequana
(Dark Spruce Bell) ws: 11-13mm; Apr-May; Norway spruce (Picea abies), silver/giant fur (Abies alba/grandis); pRDB local in S.England |
248 Epinotia tenerana
(Nut-bud Moth) 
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249 Epinotia ramella
(Small Birch Bell) 
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250 Epinotia nigricana
(Fir Bell) ws: 11-13mm; Jul; silver fir (Abies alba); pRDB coniferous woods in England |
251 Epinotia rubiginosana
(Pine Bell) 
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252 Epinotia tedella
(Common Spruce Bell) 
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253 Epinotia fraternana
(Silver-barred Bell) ws: 10-13mm; Jun-Jul; silver/giant fir (Abies alba.grandis); NS-B in coniferous woodland throughout GB |
254 Epinotia bilunana
(Crescent Bell) 
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255 Epinotia nisella
(Grey Poplar Bell) 
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256 Epinotia cinereana
(Grey Aspen Bell) 
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Zeiraphera (4S)
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257 Zeiraphera griseana
(Larch Tortrix) 
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258 Zeiraphera rufimitrana
(Red-headed Bell) ws: 12-16mm; Jul; silver fir (Abies alba), stone pine (Pinus pinea); NS-A pine woodland in S.England |
259 Zeiraphera ratzeburgiana
(Spruce-bud Moth) 
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260 Zeiraphera isertana
(Cock's-head Bell) 
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Crocidosema
261 Crocidosema plebejana
(Southern Bell) 
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Phaneta
262 Phaneta pauperana
(Scarce Rose Bell) ws: 13-17mm; Apr-May; dog rose (Rosa canina); pRDB chalky soils and sea cliffs in SE England |
Pelochrista
263 Pelochrista caecimaculana
(Chalk-hill Bell) 
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Eucosma (14S)
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264 Eucosma obumbratana
(Two-coloured Ball) 
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265 Eucosma cana
(Hoary Bell) 
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266 Eucosma hohenwartiana
(Bright Bell) 
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267 Eucosma fulvana
(Fulvous Bell) 
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268 Eucosma parvulana
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269 Eucosma campoliliana
(Marbled Bell) 
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270 Eucosma aemulana
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271 Eucosma rubescana
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272 Eucosma tripoliana
(Saltmarsh Bell) 
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273 Eucosma lacteana
(Maritime Bell) ws: 13-17mm; Jul-Aug; sea wormwood (Artemisia maritima); pRDB saltmarsh Sussex to Lincolnshire |
274 Eucosma metzneriana
(Mugwort Bell) 
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275 Eucosma conterminana
(Pale Lettuce Bell) 
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276 Eucosma aspidiscana
(Golden-rod Bell) 
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277 Eucosma pupillana
ws: 15-19mm; Jul-Aug; wormwood (Artemisia absinthium); NS-B dry pastures and waste ground in England & Wales |
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Gypsonoma (6S 1EX)
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278 Gypsonoma minutana
(Brindled Shoot) 
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279 Gypsonoma dealbana
(Common Cloaked Shoot) 
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280 Gypsonoma oppressana
(Poplar Shoot) 
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281 Gypsonoma sociana
(White Cloaked Shoot) 
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282 Gypsonoma nitidulana
Formerly at 200-300m around Aviemore. Not recorded since 1908. Extinct in GB |
283 Gypsonoma aceriana
(Rosy Cloaked Shoot) 
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Epiblema (8S)
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284 Epiblema sticticana
(Colt's-foot Bell) 
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285 Epiblema scutulana
(Thistle Bell) 
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286 Epiblema cirsiana
(Knapweed Bell) 
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287 Epiblema cnicicolana
(Fleabane Bell) ws: 14-16mm; May-Jun; common fleabane (Pulicaria dysenterica); pRDB grassland, river banks S.England & S.Wales |
288 Epiblema foenella
(White-foot Bell) 
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289 Epiblema costipunctana
(Ragwort Bell) 
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290 Epiblema turbidana
(Butterbur Bell) ws: 16-22mm; Jun-Jul; butterbur (Petasites hybridus); NS-B damp pastures and river banks throughout GB |
291 Epiblema grandaevana
(Great Bell) 21-30mm; Jul; RDB formerly established in coastal NE.England now very rare in Leicestershire and Norfolk |
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Notocelia (7S)
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292 Notocelia cynosbatella
(Yellow-faced Bell) 
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293 Notocelia tetragonana
(Square-spot Bell) ws: 13-16mm; Jul; rose (Rosa spp); NS-B open woodlands and hedgerows throughout GB, declining |
294 Notocelia uddmanniana
(Bramble Shoot Moth) 
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295 Notocelia roborana
(Summer Rose Bell) 
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296 Notocelia incarnatana
(Chalk Rose Bell) ws: 14-19mm; Jun-Sep; rose (Rosa spp); NS-B coastal limestone and sand-dunes and chalk downs |
297 Notocelia rosaecolana
(Common Rose Bell) 
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298 Notocelia trimaculana
(Triple-blotched Bell) 
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Pseudococcyx (2S)
299 Pseudococcyx posticana
(Dark Pine Shoot) 
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300 Pseudococcyx turionella
(Pine Bud Moth) 
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Retinia
301 Retinia resinella
(Pine Resin Gall Moth) ws: 16-22mm; Jun; Scots pine (Pinus sylvestris); NS-A throughout GB, most common in Scotland |
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Gravitarmata
302 Gravitarmata margarotana
(Pine Cone Tortrix) ws: 14-18mm; Apr-May; pine, fir, spruce; First British record May 2011, Clowes Wood, Kent; subsequently there have been a few records from Leicestershire, Suffolk and Norfolk |
Clavigesta (2S)
303 Clavigesta sylvestrana
(Small Pine Shoot) ws: 12-15mm; Jun-Jul; maritime/stone pine (Pinus pinaster/pinea); NS-A pine woods in S.England |
304 Clavigesta purdeyi
(Pine Leaf-mining Moth) 
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Rhyacionia (4S)
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305 Rhyacionia buoliana
(Pine Shoot) 
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306 Rhyacionia pinicolana
(Orange-spotted Shoot) 
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307 Rhyacionia pinivorana
(Spotted Shoot) 
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308 Rhyacionia logaea
(Elgin Shoot) ws: 14-18mm; Apr-May; pines (Pinus spp), Sitka spruce (Picea stichensis); NS-B pine woodland in NE.Scotland |
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