Core Stigmella - lapponica group
002 S.lapponica and 003 S.confusella are characterised by a very distinctive gnathos in which the pair of posteriorly-directed horns seen in most Stigmella species is replaced by a very strong long broad ~triangular median lobe. They differ in the following features:
- S.lapponica: uncus with a narrow median notch; aedeagus with ~70 small thorns the apical ~20 of which are distinctly larger than the rest
- S.confusella: uncus with a broad median notch: aedeagus with ~70 small thorns without a size contrast
013 S.malella is included in the lapponica group but is clearly something of an imposter, having a more standard gnathos with a pair of horns. The only feature that appears to unify the 3 species is the presence of an arcuate process at the apex of the valva. Other features of S.malella are: uncus bilobed with a U-shaped median notch; aedeagus slightly shorter than genital capsule with two connected bundles of elongate triangular spines, the apical bundle containing slightly larger spines than the anterior bundle.
Non-core Stigmella - tiliae group
004 S.tiliae only. Characterised by the cornuti consisting of an anterior cordate plate with two long needle-like cornuti extending posteriorly from each lateral margin of the plate.
Non-core Stigmella - betulicola group
The 6 species in this group are characterised by a very short aedeagus, ½ or <½ the length of the genital capsule, with cornuti in the form of 2 crenate plates at the apex of the aedeagus and a small anterior bundle of spines; they also all have widely separated fairly long gnathos horns which curve away from the midline. the species differ mainly in the shapes of 1) the hind margin of the uncus, 2) the anterior emargination of the vinculum and 3) the apex of the valvae.
- 005 S.betulicola - the only member of the group with distinct lateral processes of the transtilla and the only member of the group with narrow anterior angles of the vinculum. 1) ~straight; 2) moderate U; 3) tapered apical ¼
- 006 S.sakhalinella - 1) concave; 2) shallow; 3) abruptly narrowed
- 007 S.luteella - 1) concave: 2) deep U; 3) tapered apical ⅙
- 008 S.glutinosae - 1) ~straight; 2) deep V; 3) tapered apical ⅙
- 009 S.alnetella - 1) broad angular emargination with distinct lateral processes; 2) deep V; 3) tapered apical ½
- 010 S.microtheriella - no information
Non-core Stigmella - prunifoliella group
011 S.prunetorum only. I have not found any images of the genitalia and it is not clear to me what feature justifies its placement in a separate group
Vinculum with a deep anterior emargination and pointed lateral angles; uncus short and broad with a shallowly concave hind margin and sclerotised lateral margins (said to be X-shaped, but I am not sure what this means); ganthos with long narrow horizontal bar, long narrow horns and short anterior processes; valvae slender, arcuate; transtilla with long lateral processes; juxta triangular; aedeagus ¾-⅘ length of capsule containing many small cornuti.
Vinculum with a deep anterior emargination and pointed lateral angles; uncus short and broad with a shallowly concave hind margin and sclerotised lateral margins (said to be X-shaped, but I am not sure what this means); ganthos with long narrow horizontal bar, long narrow horns and short anterior processes; valvae slender, arcuate; transtilla with long lateral processes; juxta triangular; aedeagus ¾-⅘ length of capsule containing many small cornuti.
Non-core Stigmella - ultima group
012 S.aceris only. Characterised by bifid valval apex and very long aedeagus (1.3-1.5x length of genital capsule). Other features: vinculum with broad U-shaped anterior emargination and pointed lateral angles; uncus with shallow emargination of posterior margin and sclerotised lateral margins; gnathos with long straight horizontal bar, long widely spaced horns and distinct anterior processes; transtilla with long lateral processes; vesica containing numerous very small cornuti.
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Core Stigmella - rhamnella group
014 S.catharticella only. Characterised by the aedaegus with 30-35 large triangular cornuti, a patch of smaller thinner apical spines and a cathrema with a well-sclerotised curved posterior extension. Other features: vinculum with a shallow anterior emargination and rounded lateral angles; uncus with small projections and sclerotised lateral margins; gnathos with a broad bar and short horns; valva short; transtilla with ~triangular lateral processes; aedaegus slightly shorter than genital capsule
Core Stigmella - anomalella group
The 3 species in this group are characterised by a bilobed (M-shaped) uncus with the median emargination about the same size as an uncus lobe; a gnathos with a short transverse bar and long relatively broad horns an a U-shaped transtilla without lateral processes. The species differ mainly features of the aedeagus.
- 015 S.anomalella - aedeagus slightly short than genital capsule; 6-8 pointed small cornuti and sparse smaller cornuti
- 016 S.spinosissimae - aedeagus slightly shorter than genital capsule; 2 apical groups of numerous long slender cornuti and a patch of small spines
- 017 S.centifoliella - aedeagus distinctly shorter than genital capsule; 2 or 3 groups of blunt small cornuti and numerous smaller spines in 2 groups (more easily distinguished on external features as it is the only member of the group with a forewing fascia)
Non-core Stigmella - ulmivora group
The 2 species in this group seem to have quite different genitalia; the only unifying feature I can determine is a bilobed uncus truncate lobes.
018 S.ulmivora. vinculum with broad U-shaped anterior emargination and long pointed lateral angles; gnathos with a moderate transverse bar, horns and anterior processes; valvae triangular, arcuate; transtilla with long triangular lateral processes; aedeagus as long as capsule densely packed with large triangular spines and dentate plates.
019 S.viscerella. vinculum with broad U-shaped anterior emargination and short pointed lateral angles; gnathos with a broad rectangular transverse bar, and robust horns; valvae triangular, arcuate, with a distinct inner lobe; transtilla with blunt lateral processes; aedeagus ½ as long as capsule with 25-35 small spines and 50+ smaller ones and a wide prominent cathrema.
018 S.ulmivora. vinculum with broad U-shaped anterior emargination and long pointed lateral angles; gnathos with a moderate transverse bar, horns and anterior processes; valvae triangular, arcuate; transtilla with long triangular lateral processes; aedeagus as long as capsule densely packed with large triangular spines and dentate plates.
019 S.viscerella. vinculum with broad U-shaped anterior emargination and short pointed lateral angles; gnathos with a broad rectangular transverse bar, and robust horns; valvae triangular, arcuate, with a distinct inner lobe; transtilla with blunt lateral processes; aedeagus ½ as long as capsule with 25-35 small spines and 50+ smaller ones and a wide prominent cathrema.
Core Stigmella - oxyacanthella group
The 9 species in this group are characterised by a bilobed uncus, a vinculum with a ~semicircular anterior emargination and pointed lateral angles, transtilla with short triangular lateral processes, and a long aedaegus packed with small cornuti. 2 of these species are regarded as extinct in GB and 1 (S.magdalenae) may be a variant of S.nylandriella. Species differ in details of the uncus lobes and emargination between them, form of the gnathos, length of the distal process of the valva, shape of the posterior margin of the inner lobe of the valva, length of aedeagus and form of cornuti. This is a difficult group due to variability within each species and similarity between the species
021 S.torminalis (extinct) - aedeagus 1.5x length of genital capsule, some cornuti relatively large and pointed; gnathos with a short transverse bar and moderate horns
022 S.regiella - aedeagus slightly longer than genital capsule, most cornuti triangular, larger cornuti more pointed; valva with posterior margin of inner lobe straight and at ~90° to distal process; gnathos short with moderate robust horns
023 S.crataegella - aedeagus ~1.3 x genital capsule; valva with posterior margin of inner lobe angular or rounded; gnathos moderate with short robust horns.
(024 S.magdalenae)
025 S.nylandriella - aedeagus 1.1 - 1.3x genital capsule; cornuti variable semicircular, obtuse or sharply-pointed; uncus lobes small and widely separated; gnathos variable, usually with a long transverse bar and short horns; valva with rounded inner lobe. Distal process of valva slightly longer than in S.oxacanthella.
026 S.oxyacanthella - aedeagus slightly shorter than genital capsule, cornuti truncate or blunt; posterior margin of uncus shallowly concave; gnathos with a moderate transverse bar, short robust horns and long pointed anterior processes; inner lobe of valva angular. Distal process of valva slightly shorter than in S.nylandriella
027 S.pyri - aedeagus slightly longer than genital capsule; posterior margin of uncus with an angular emargination so that the lateral lobes are more prominent than in S.oxyacanthella. Distal process of valva slightly longer than in S.nylandriella
028 S.minisculella - aedeagus 1.2 - 1.4x genital capsule, cornuti small and circular at anterior end becoming needle-shaped near apex. Valva with very long distal process.
029 S.desperatella (extinct) - aedeagus ~same length as genital capsule, cornuti oval to triangular; gnathos with short transverse bar and short horns; distal process of valva long extending posterior to uncus, inner lobe of valva pointed.
I will do more work on this group once I have obtained some material.
021 S.torminalis (extinct) - aedeagus 1.5x length of genital capsule, some cornuti relatively large and pointed; gnathos with a short transverse bar and moderate horns
022 S.regiella - aedeagus slightly longer than genital capsule, most cornuti triangular, larger cornuti more pointed; valva with posterior margin of inner lobe straight and at ~90° to distal process; gnathos short with moderate robust horns
023 S.crataegella - aedeagus ~1.3 x genital capsule; valva with posterior margin of inner lobe angular or rounded; gnathos moderate with short robust horns.
(024 S.magdalenae)
025 S.nylandriella - aedeagus 1.1 - 1.3x genital capsule; cornuti variable semicircular, obtuse or sharply-pointed; uncus lobes small and widely separated; gnathos variable, usually with a long transverse bar and short horns; valva with rounded inner lobe. Distal process of valva slightly longer than in S.oxacanthella.
026 S.oxyacanthella - aedeagus slightly shorter than genital capsule, cornuti truncate or blunt; posterior margin of uncus shallowly concave; gnathos with a moderate transverse bar, short robust horns and long pointed anterior processes; inner lobe of valva angular. Distal process of valva slightly shorter than in S.nylandriella
027 S.pyri - aedeagus slightly longer than genital capsule; posterior margin of uncus with an angular emargination so that the lateral lobes are more prominent than in S.oxyacanthella. Distal process of valva slightly longer than in S.nylandriella
028 S.minisculella - aedeagus 1.2 - 1.4x genital capsule, cornuti small and circular at anterior end becoming needle-shaped near apex. Valva with very long distal process.
029 S.desperatella (extinct) - aedeagus ~same length as genital capsule, cornuti oval to triangular; gnathos with short transverse bar and short horns; distal process of valva long extending posterior to uncus, inner lobe of valva pointed.
I will do more work on this group once I have obtained some material.
Core Stigmella - hybnerella group
The 3 members of this group are characterised by a very broad uncus with little in the way of lateral processes; a gnathos with a very long transverse bar and widely spaced horns; a very short distal process of the valva; transtilla with a long transverse bar. The vinculum is broadly emarginate with some intraspecific variation in the depth of the excavation. The species differ in overall shape of the genital capsule, shape of the anterior margin of the vinculum
020 S.paradoxa - genital capsule almost globular; vinculum with blunt lateral angles; inner lobe of valva large, rounded, almost reaching tip of distal process of valva; transtilla without lateral processes; aedeagus as long as capsule, packed with spines and curved thorns.
030 S.hybnerella- genital capsule longer than wide; vinculum with moderate pointed lateral angles; inner lobe of valva large, rounded, reaching hind margin of uncus; transtilla with moderate slender lateral processes; aedeagus 1.4 - 1.6x genital capsule, with a long band of hundreds of small spines and 3-6 triangular cornuti at apex.
031 S.mespilicola - genital capsule as long as wide; vinculum with short pointed lateral angles; inner lobe of valva short, blunt, not reaching hind margin of uncus; transtilla without lateral processes; aedeagus longer than genital capsule, with a long band of hundreds of small spines without triangular cornuti at apex.
020 S.paradoxa - genital capsule almost globular; vinculum with blunt lateral angles; inner lobe of valva large, rounded, almost reaching tip of distal process of valva; transtilla without lateral processes; aedeagus as long as capsule, packed with spines and curved thorns.
030 S.hybnerella- genital capsule longer than wide; vinculum with moderate pointed lateral angles; inner lobe of valva large, rounded, reaching hind margin of uncus; transtilla with moderate slender lateral processes; aedeagus 1.4 - 1.6x genital capsule, with a long band of hundreds of small spines and 3-6 triangular cornuti at apex.
031 S.mespilicola - genital capsule as long as wide; vinculum with short pointed lateral angles; inner lobe of valva short, blunt, not reaching hind margin of uncus; transtilla without lateral processes; aedeagus longer than genital capsule, with a long band of hundreds of small spines without triangular cornuti at apex.
Core Stigmella - floslactella group
The 3 members of this group are characterised by a bilobed uncus, the lobes separated by an angular emargination, and each lobe bifid to some degree; a gnathos with long slender horns; a vinculum with a moderate to deep anterior emargination
032 S.floslactella - aedeagus as long as genital capsule, with an anterior group of needle-like cornuti and an apical group of ~25 broad short cornuti; uncus lobes shallowly bifid
033 S.carpinella - aedeagus shorter than genital capsule, with 30-35 broad cornuti and numerous weakly sclerotised small spines; uncus lobes shallowly bifid
034 S.tityrella - aedeagus as long as genital capsule, with an anterior group of 10-20 small triangular cornuti and an apical group of ~20 larger triangular cornuti; uncus lobes distinctly bifid giving the hind margin of the uncus as a whole a dentate appearance
032 S.floslactella - aedeagus as long as genital capsule, with an anterior group of needle-like cornuti and an apical group of ~25 broad short cornuti; uncus lobes shallowly bifid
033 S.carpinella - aedeagus shorter than genital capsule, with 30-35 broad cornuti and numerous weakly sclerotised small spines; uncus lobes shallowly bifid
034 S.tityrella - aedeagus as long as genital capsule, with an anterior group of 10-20 small triangular cornuti and an apical group of ~20 larger triangular cornuti; uncus lobes distinctly bifid giving the hind margin of the uncus as a whole a dentate appearance
Core Stigmella - salicis group
The 6 members of this group are characterised by an aedeagus with a relatively small number of relatively large separate cornuti in the apical ~½ and a cluster of spines in the basal ½; and a gnathos with lateral arms as well as horns. Uncus bilobed, the lobes bifid to a variable degree. The transtilla of most species has short triangular lateral processes. The male genitalia S.salicis and S.myrtilella are extremely similar to each other and can only be separated on details of the cornuti; others species also differ in the arrangement of the cornuti but can also be distinguished on details of the uncus, gnathos and posterior margin of the valva.
035 S.salicis - aedeagus slightly shorter than capsule, cornuti 4-8 (usually 5 or 6) of unequal length and ~straight and a small anterior cluster of 4-8 slender spines of unequal length; median notch of uncus deep and angular, lobes bifid to a variable degree; gnathos horns separate, parallel to slightly diverging, lateral arms short, narrow; posterior margin of valva with two pointed process (one on distal process other on inner lobe, incurved) 036 S.myrtilella - aedeagus slightly shorter than capsule, cornuti 3-7 (usually 3 or 4) of equal length, short and ~straight and a small anterior cluster of 4-7 slender spines of equal length; median notch of uncus deep and angular, lobes distinctly bifid; gnathos horns close-set at base, slightly diverging, lateral arms short, narrow; posterior margin of valva with two pointed process (one on distal process other on inner lobe, incurved) 037 S.zelleriella - aedeagus distinctly shorter than capsule, cornuti 5-6 unequal and a small anterior cluster of 5-9 spines of unequal length; median notch of uncus deep and angular, lobes shallowly bifid; gnathos horns separate at base, parallel, lateral arms "distinct"; posterior margin of valva with two short pointed process (one on distal process other on inner lobe, incurved) 038 S.obliquella - aedeagus distinctly shorter than capsule, cornuti 6-7 of unequal length and ~conical and a small anterior cluster of 2-3 long (curved?) slender spines; median notch of uncus deep and narrow, lobes shallowly bifid; gnathos horns fused at base, parallel, lateral arms short, broad; posterior margin of valva with a pointed distal process and a rounded inner lobe. Lateral angles of vinculum and lateral processes of transtilla broader and blunter than in other species in the group 039 S.trimaculella - aedeagus distinctly shorter than capsule, cornuti 3-4 small and an anterior cluster of 8-10 straight slender spines; median notch of uncus deep V, lobes indistinctly or not bifid; gnathos horns close-set at base, slightly diverging, lateral arms short, narrow; posterior margin of valva with two short pointed process (one on distal process other on inner lobe). Anterior emargination of vinculum deeper than in other members of the group. Transtilla with a longer transverse bar and narrower, less triangular lateral processes than in other members of the group. 040 S.assimilella - aedeagus slightly shorter than capsule, cornuti 5-6 and an anterior cluster of 10-15 sharply pointed spines some of which are curved; median notch of uncus deep angular, lobes shallowly bifid; gnathos horns close-set at base, slightly diverging, lateral arms moderate, narrow; posterior margin of valva with tapered distal process and straight or angular inner lobe. Anterior emargination of vinculum shallower than in other members of the group. Transtilla with a broad transverse bar and short blunt lateral processes. |
Core Stigmella - sorbi group
The 2 members of this group are characterised by a distinctly elongate genital capsule, this appearance being largely due to a long narrow vinculum with a narrow deep anterior emargination; uncus bilobed, each lobe bifid; valva with two apical processes; aedeagus as long as capsule
041 S.sorbi - aedagus packed with many needle-like cornuti, apex with a sclerotised plate and 8-10 strong cornuti; gnathos horns widely spaced parallel; manica sparsely spined
042 S.plagicolella - aedagus packed with spines of various sizes, apex with a sclerotised plate and a few large cornuti; gnathos horns close-set diverging (forming a V); manica large, distinctly spined
041 S.sorbi - aedagus packed with many needle-like cornuti, apex with a sclerotised plate and 8-10 strong cornuti; gnathos horns widely spaced parallel; manica sparsely spined
042 S.plagicolella - aedagus packed with spines of various sizes, apex with a sclerotised plate and a few large cornuti; gnathos horns close-set diverging (forming a V); manica large, distinctly spined
Core Stigmella - lemniscella group
The 2 members of this group are characterised by a vinculum with a deep anterior emargination a small bilobed uncus, each lobe bifid; a bulb-shaped aedeagus about as long as the capsule, packed with small spines
043 S.lemniscella - gnathos horns forming a V; apical cornuti blunt; lateral processes of transtilla blunt; manica sparsely spined
044 S.continuella - gnathos horns parallel; apical cornuti triangular; lateral processes of transtilla pointed; manica distinctly spined
The 2 members of this group are characterised by a vinculum with a deep anterior emargination a small bilobed uncus, each lobe bifid; a bulb-shaped aedeagus about as long as the capsule, packed with small spines
043 S.lemniscella - gnathos horns forming a V; apical cornuti blunt; lateral processes of transtilla blunt; manica sparsely spined
044 S.continuella - gnathos horns parallel; apical cornuti triangular; lateral processes of transtilla pointed; manica distinctly spined
Core Stigmella - aurella group
(Only S.aurella and S.dryadella are shown at Moth Dissection)
The 8 members of this group have a vinculum with a moderate anterior emargination; a bilobed uncus and usually shallowly bifid lobes; gnathos with separate parallel horns; a broad and short valva; a transtilla with blunt but usually distinct short lateral processes; aedeagus bulbed. The species differ particularly in details of the aedeagus and distal process of the valva.
045 S.aurella - valva with a pointed distal process at 90° to the outer margin; aedeagus 1.5x length of genital capsule, packed with numerous needle-shaped cornuti and with a few larger conical cornuti at the apex
046 S.auromarginella - valva with a short triangular distal process; aedeagus slightly longer than capsule with an irregular (boot-shaped) cornutus near apex
047 S.splendidissimella - anterior emargination of vinculum narrow and lateral angles broad; valva constricted "before middle", distal process long and slightly curved; transtilla very broad, lateral processes indistinct; aedeagus slightly longer than capsule with numerous small cornuti and small spines
048 S.pretiosa - anterior emargination of vinculum narrow and lateral angles broad; valva constricted "before middle", distal process triangular, at 90° to outer margin; transtilla broad; aedeagus as long as capsule with 7-12 pointed cornuti near apex
049 S.aeneofasciella - anterior emargination of vinculum shallow; uncus very broad, lobes distinctly bifid; distal process of valva short and blunt; aedeagus as long as capsule with 3-4 strong broad-based cornuti and some small spines near apex, and an anterior group of ~15 close-set strong cornuti.
050 S.dryadella - anterior emargination of vinculum shallow; uncus lobes distinctly bifid; distal process of valva short, pointed, at 90° to outer margin; aedeagus slightly longer than capsule with one long curved broad-based cornutus near apex and an anterior bundle of ~30 varied cornuti
051 S.poterii - uncus large with deep narrow median notch, the lobes not (or very slightly) bifid; distal process of valva long and blunt; aedeagus slightly longer than capsule with one long curved broad-based cornutus near apex and an anterior bundle of ~100 needle-like cornuti
052 S.filipendulae - uncus small with U-shaped medial notch, the lobes distinctly bifid; distal process of valva strong, pointed; aedeagus slightly shorter than capsule with one long curved broad-based cornutus near apex and an anterior bundle of ~100 needle-like cornuti
(Only S.aurella and S.dryadella are shown at Moth Dissection)
The 8 members of this group have a vinculum with a moderate anterior emargination; a bilobed uncus and usually shallowly bifid lobes; gnathos with separate parallel horns; a broad and short valva; a transtilla with blunt but usually distinct short lateral processes; aedeagus bulbed. The species differ particularly in details of the aedeagus and distal process of the valva.
045 S.aurella - valva with a pointed distal process at 90° to the outer margin; aedeagus 1.5x length of genital capsule, packed with numerous needle-shaped cornuti and with a few larger conical cornuti at the apex
046 S.auromarginella - valva with a short triangular distal process; aedeagus slightly longer than capsule with an irregular (boot-shaped) cornutus near apex
047 S.splendidissimella - anterior emargination of vinculum narrow and lateral angles broad; valva constricted "before middle", distal process long and slightly curved; transtilla very broad, lateral processes indistinct; aedeagus slightly longer than capsule with numerous small cornuti and small spines
048 S.pretiosa - anterior emargination of vinculum narrow and lateral angles broad; valva constricted "before middle", distal process triangular, at 90° to outer margin; transtilla broad; aedeagus as long as capsule with 7-12 pointed cornuti near apex
049 S.aeneofasciella - anterior emargination of vinculum shallow; uncus very broad, lobes distinctly bifid; distal process of valva short and blunt; aedeagus as long as capsule with 3-4 strong broad-based cornuti and some small spines near apex, and an anterior group of ~15 close-set strong cornuti.
050 S.dryadella - anterior emargination of vinculum shallow; uncus lobes distinctly bifid; distal process of valva short, pointed, at 90° to outer margin; aedeagus slightly longer than capsule with one long curved broad-based cornutus near apex and an anterior bundle of ~30 varied cornuti
051 S.poterii - uncus large with deep narrow median notch, the lobes not (or very slightly) bifid; distal process of valva long and blunt; aedeagus slightly longer than capsule with one long curved broad-based cornutus near apex and an anterior bundle of ~100 needle-like cornuti
052 S.filipendulae - uncus small with U-shaped medial notch, the lobes distinctly bifid; distal process of valva strong, pointed; aedeagus slightly shorter than capsule with one long curved broad-based cornutus near apex and an anterior bundle of ~100 needle-like cornuti
Core Stigmella - incognitella group
The 2 species in this group are characterised by a distinctly elongate genital capsule with a long vinculum with a very deep anterior emargination and long blunt lateral processes; uncus bilobed, the lobes shallowly bifid; valva with a short blunt distal process; transtilla with long lateral processes; aedaegus somewhat shorter than capsule, cylindrical with 2 groups of cornuti, one with 30-40 small spines the other with ~100 very small spines.
053 S.incognitella - gnathos bar + horns V-shaped, with broad lateral arms
054 S.perpygmaeella - gnathos bar + horns U-shaped, with moderate lateral arms
053 S.incognitella - gnathos bar + horns V-shaped, with broad lateral arms
054 S.perpygmaeella - gnathos bar + horns U-shaped, with moderate lateral arms
Core Stigmella - ruficapitella group
The 8 species of this group are characterised by the uncus with posterolateral processes widely separated by a broad concavity of the hind margin; the gnathos horns are widely spaced in all except S.hemargyrella; transtilla a curved bar parallel to the anterior margin of the vinculum
055 S.hemargyrella - uncus with a median hump; distal lobe of valva short; gnathos horns close together; aedeagus longer than capsule, packed with numerous conical cornuti the largest near the apex
057 S.suberivora - genital capsule at least as wide as long with bulging valvae; "vinculum anteriorly slightly bilobed"?; gnathos horns widely spaced sometimes converging posteriorly; distal <⅓ of valva forming a pointed process; aedeagus with an anterior bundle of 20-30 curved cornuti with serrate bases together with many smaller spines and an apical group of 10-12 long needles and one or more larger cornuti at the apex; manica almost completely covering aedeagus.
058 S.basiguttella - aedeagus almost 2x length of genital capsule, with a large bundle of long curved spines occupying most of the vesica and a smaller apical bundle of smaller spines with 1 or a few larger cornuti at the apex
059 S.svenssoni - aedeagus slightly longer than capsule, with a large bundle of long curved spines occupying most of the vesica and a smaller apical bundle of smaller spines
060 S.ruficapitella - manica conspicuous, almost 2x width of aedeagus; aedeagus slightly longer than capsule and packed with mostly needle-like spines
061 S.atricapitella - manica inconspicuous, narrow; aedeagus longer than capsule, packed cornuti of various sizes including many long broad-based tapering spines.
062 S.samiatella - aedagus ~1.2x length of capsule, with a much lower density of cornuti than others in the group, arranged in 3 well separated bundles: an anterior plate with several short robust spines, a narrow central line of small spines becoming longer towards the apex, and an apical group of broader based more robust short cones.
063 S.roborella - posterior margin of valva with two points - the distal process and a pointed inner lobe; aedeagus with cornuti arranged in 3 bundles without much separation between them: a dense anterior group of medium spines, a central group of more robust cones and a smaller apical group of smaller spines.
055 S.hemargyrella - uncus with a median hump; distal lobe of valva short; gnathos horns close together; aedeagus longer than capsule, packed with numerous conical cornuti the largest near the apex
057 S.suberivora - genital capsule at least as wide as long with bulging valvae; "vinculum anteriorly slightly bilobed"?; gnathos horns widely spaced sometimes converging posteriorly; distal <⅓ of valva forming a pointed process; aedeagus with an anterior bundle of 20-30 curved cornuti with serrate bases together with many smaller spines and an apical group of 10-12 long needles and one or more larger cornuti at the apex; manica almost completely covering aedeagus.
058 S.basiguttella - aedeagus almost 2x length of genital capsule, with a large bundle of long curved spines occupying most of the vesica and a smaller apical bundle of smaller spines with 1 or a few larger cornuti at the apex
059 S.svenssoni - aedeagus slightly longer than capsule, with a large bundle of long curved spines occupying most of the vesica and a smaller apical bundle of smaller spines
060 S.ruficapitella - manica conspicuous, almost 2x width of aedeagus; aedeagus slightly longer than capsule and packed with mostly needle-like spines
061 S.atricapitella - manica inconspicuous, narrow; aedeagus longer than capsule, packed cornuti of various sizes including many long broad-based tapering spines.
062 S.samiatella - aedagus ~1.2x length of capsule, with a much lower density of cornuti than others in the group, arranged in 3 well separated bundles: an anterior plate with several short robust spines, a narrow central line of small spines becoming longer towards the apex, and an apical group of broader based more robust short cones.
063 S.roborella - posterior margin of valva with two points - the distal process and a pointed inner lobe; aedeagus with cornuti arranged in 3 bundles without much separation between them: a dense anterior group of medium spines, a central group of more robust cones and a smaller apical group of smaller spines.
Core Stigmella - speciosa group
056 S.speciosa - vinculum with a shallow anterior emargination and rounded lateral angles; uncus bilobed, the lobes bifid; gnathos with fairly short parallel horns; distal process of valva very long, straight and narrow, inner lobe short; transtilla with slender transverse bar and triangular lateral processes; aedeagus much longer than capsule with a large chitin plate at apex and packed with triangular cornuti that graduate from small to larger towards the apex.